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The first interchangeable lenses, as we know, were the non-standardized lenses for the early (2995 made) Ic non-standard cameras. They carried the whole or partial serial number of the camera on the lens depth of field ring. By 1931 the lens flange to film distance was standardized, and these new lenses were marked with a small "o", again on the depth ring. Early lenses could be returned, with camera, for "standardization", and the "o" was added but the match number was normally not removed. With the non-standard lenses, Leitz had the 35 Elmar, 50 Elmar, 50 Hektor and the 135 Elmar. Before Rangefinder coupling, before lens serialization, before infinity lock devices, before the 47mm flange, 11 O'clock standardized lenses were produced with the "o" mark. I believe the earliest of these lenses had an "o" on both the depth ring and also on the rear of the lens flange. Here is photo of 5 of the first lenses, all uncoupled, no serials, no infinity locks. The next photo shows a close up of the 35, and 50's back flange with the additional "o", ( look closely). From my checks, this second "o" was not used long, and does not appear on lens with serials,or infinity locks. The 135, added the "o" to the arrow at the base, but it is not on the rear of the flange. The 90 Elmar was not introduced until the standardized camera and is not seen with the compatibility number, or "o" mark, but is sometimes uncoupled as as the coupled camera was not until Feb. 1932. So, all 90 Elmars are standardized from the first, my theory. Any additional observations of the rear "o" is welcome, check your early lenses. Any corrections also most welcome. Stay safe.